In the present day, two years after the events of the previous film, Optimus Prime is seen leading NEST, a military organization consisting of human troops and his own team of Autobots (including newcomers Arcee, Chromia, Elita One, Sideswipe, Jolt, and the twins Skids and Mudflap) aimed at killing the remaining Decepticons on Earth. The remaining brothers thus sacrificed their bodies in order to hide the Matrix of Leadership the key that activates the Sun Harvester from The Fallen, who swore to seek revenge upon Earth. The Primes agreed that life-bearing worlds would be spared, but in 17,000 BC, one brother, thereafter dubbed "The Fallen", constructed a Sun Harvester on Earth. Known as the Dynasty of Primes, they used machines called Sun Harvesters to drain stars of their energy in order to convert it to energon and power Cybertron's AllSpark. It is revealed that thousands of years ago there was a race of ancient Transformers who scoured the universe looking for energon sources.
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